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The cause of the mishap came down to fatigue and bad communication. A fuel leak had developed in the accident wing in the #2 main tank. The plane was pulled into the fuel barn (the hangar in the background of the photos). The leak was repaired and pressure plugs were inserted in the place of the vent plugs. The wing was charged with compressed air. The theory being that if the tank would hold a certain pressure for a certain time, then the leak must be fixed. This also allowed for the replaced panels to "seat."
The plane was then taken back out to the line and refueling started for her next trip to Germany later that day. The personnel who performed the repairs had put in a 20 hour day and went home. Follow on personnel did not swap the pressure test plugs for the vent plugs. As fuel flowed into the #2 main, which is the closest to the fuselage, the air had nowhere to go and compressed until it blew out the skin. Unfortunately, the pressure was so great that it broke the wing spar in the process.